Defensive alliances in spatial models of cyclical population interactions

As a generalization of the three-strategy Rock-Scissors-Paper game dynamics in space, cyclical interaction models of six mutating species are studied on a square lattice, in which each species is supposed to have two dominant, two subordinated, and a neutral interacting partner. Depending on their interaction topologies, all imaginable systems can be classified into four (isomorphic) groups exhibiting significantly different behaviors as a function of mutation rate. In three out of four cases three (or four) species form defensive alliances that maintain themselves in a self-organizing polydomain structure via cyclic invasions. Varying the mutation rate, this mechanism results in an ordering phenomenon analogous to that of magnetic Ising systems. The model explains a very basic mechanism of community organization, which might gain important applications in biology, economics, and sociology.