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CDKN1C/p57kip2 is a candidate tumor suppressor gene in human breast cancer
Pamela S Larson1, Benjamin L Schlechter, Chia-Lin King
1Department of Pathology and Laboratory Medicine, Boston University Medical Center, Boston, MA, USA. pamkent@bu.edu
Background:
CDKN1C (also known as p57KIP2) is a cyclin-dependent kinase inhibitor previously implicated in several types of human cancer. Its family members (CDKN1A/p21CIP1 and B/p27KIP1) have been implicated in breast cancer, but information about CDKN1C's role is limited. We hypothesized that decreased CDKN1C may be involved in human breast carcinogenesis in vivo.
Methods:
We determined rates of allele imbalance or loss of heterozygosity (AI/LOH) in CDKN1C, using an intronic polymorphism, and in the surrounding 11p15.5 region in 82 breast cancers. We examined the CDKN1C mRNA level in 10 cancers using quantitative real-time PCR (qPCR), and the CDKN1C protein level in 20 cancers using immunohistochemistry (IHC). All samples were obtained using laser microdissection. Data were analyzed using standard statistical tests.
Results:
AI/LOH at 11p15.5 occurred in 28/73 (38%) informative cancers, but CDKN1C itself underwent AI/LOH in only 3/16 (19%) cancers (p = ns). In contrast, CDKN1C mRNA levels were reduced in 9/10 (90%) cancers (p < 0.0001), ranging from 2-60% of paired normal epithelium. Similarly, CDKN1C protein staining was seen in 19/20 (95%) cases' normal epithelium but in only 7/14 (50%) cases' CIS (p < 0.004) and 5/18 (28%) cases' IC (p < 0.00003). The reduction appears primarily due to loss of CDKN1C expression from myoepithelial layer cells, which stained intensely in 17/20 (85%) normal lobules, but in 0/14 (0%) CIS (p < 0.00001). In contrast, luminal cells displayed less intense, focal staining fairly consistently across histologies. Decreased CDKN1C was not clearly associated with tumor grade, histology, ER, PR or HER2 status.
Conclusion:
CDKN1C is expressed in normal epithelium of most breast cancer cases, mainly in the myothepithelial layer. This expression decreases, at both the mRNA and protein level, in the large majority of breast cancers, and does not appear to be mediated by AI/LOH at the gene. Thus, CDKN1C may be a breast cancer tumor suppressor.
Insights
CDKN1C (p57KIP2) expression decreases in most breast cancers at both mRNA and protein levels. This reduction is not due to gene mutation, suggesting CDKN1C may act as a tumor suppressor in breast cancer.
Area of Science:
- Oncology
- Molecular Biology
- Genetics
Background:
- CDKN1C (p57KIP2) is a cyclin-dependent kinase inhibitor with limited known involvement in breast cancer.
- Its family members are implicated in breast cancer, prompting investigation into CDKN1C's role.
- Hypothesis: Decreased CDKN1C expression contributes to breast carcinogenesis.
Purpose of the Study:
- To investigate the role of CDKN1C in breast cancer development.
- To determine if CDKN1C expression is altered in breast tumors.
- To explore the mechanism of CDKN1C alteration in breast carcinogenesis.
Main Methods:
- Analyzed allele imbalance/loss of heterozygosity (AI/LOH) of CDKN1C and the 11p15.5 region in 82 breast cancers.
- Quantified CDKN1C mRNA levels using qPCR in 10 cancers.
- Assessed CDKN1C protein levels via immunohistochemistry in 20 cancers, using laser microdissection for all samples.
Main Results:
- AI/LOH at 11p15.5 occurred in 38% of cancers, but CDKN1C itself showed AI/LOH in only 19%.
- CDKN1C mRNA levels were reduced in 90% of cancers (2-60% of normal levels).
- CDKN1C protein was decreased in ductal carcinoma in situ (50%) and invasive carcinoma (28%), primarily due to loss of expression in myoepithelial cells.
Conclusions:
- CDKN1C is expressed in normal breast epithelium, predominantly in myoepithelial cells.
- A significant decrease in both mRNA and protein levels of CDKN1C occurs in most breast cancers.
- The reduction in CDKN1C is not mediated by AI/LOH, suggesting a tumor suppressor role for CDKN1C in breast cancer.
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