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Neurulation01:30

Neurulation

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Neurulation is the embryological process which forms the precursors of the central nervous system and occurs after gastrulation has established the three primary cell layers of the embryo: ectoderm, mesoderm, and endoderm. In humans, the majority of this system is formed via primary neurulation, in which the central portion of the ectoderm—originally appearing as a flat sheet of cells—folds upwards and inwards, sealing off to form a hollow neural tube. As development proceeds, the...
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Surface Appendages of Archaea01:23

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Archaeal surface appendages are highly specialized structures essential for environmental adaptation, encompassing roles in adhesion, biofilm formation, and motility. Among these appendages, pili and archaella stand out for their distinct morphologies and functionalities, enabling archaea to thrive in diverse and often extreme environments.Pili: Adhesion and Biofilm FormationPili are filamentous structures assembled from pilin protein subunits, primarily contributing to adhesion and biofilm...
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Diversity of Protists II01:27

Diversity of Protists II

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Alveolates are a group of organisms recognized by the presence of alveoli, which are cytoplasmic sacs located beneath the cell membrane. While their function remains uncertain, alveoli may help regulate water balance by controlling how much water enters and leaves the cell. In dinoflagellates, these structures may serve as armor plates. There are three major types of alveolates: ciliates, which move using cilia; dinoflagellates, which use flagella for movement; and apicomplexans, which are...
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Diversity of Protists III01:27

Diversity of Protists III

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Rhizaria are a diverse group of unicellular protists characterized by their threadlike cytoplasmic extensions known as pseudopodia. These structures aid in both locomotion and feeding, giving Rhizaria an amoeboid appearance. Their amoeboid morphology once led to taxonomic confusion, but molecular phylogenetics has clarified their evolutionary placement and emphasized their shared use of pseudopodia despite divergent lineages.This clade comprises diverse lineages such as Chlorarachniophyta,...
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Cleavage and Blastulation01:33

Cleavage and Blastulation

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After a large-single-celled zygote is produced via fertilization, the process of cleavage occurs while zygotes travel through the uterine tube. Cleavage is a mitotic cell division that does not result in growth. With each round of successive cell division, daughter cells get increasingly smaller.
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Cephalic Phase of Digestion01:24

Cephalic Phase of Digestion

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The process of digestion is composed of three stages – cephalic, gastric, and intestinal – each with a distinct control center. The cephalic phase is the first stage, and it starts even before the food enters the stomach. It is controlled by the central nervous system and is initiated by any food-related sensory stimuli, such as the sight and smell of food, which send signals to the brain. While eating, the taste receptors intensify these signals, which travel to the cerebral cortex...
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Related Experiment Video

Updated: Nov 22, 2025

Genotyping of Sea Anemone during Early Development
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Genotyping of Sea Anemone during Early Development

Published on: May 13, 2019

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Primitive soft-bodied cephalopods from the Cambrian.

Martin R Smith1, Jean-Bernard Caron

  • 1Department of Ecology and Evolutionary Biology, University of Toronto, 25 Harbord Street, Toronto, Ontario, M5S 3G5, Canada.

Nature
|May 28, 2010
PubMed
Summary

Early cephalopods, like Nectocaris, lacked shells and lived in the Cambrian period. This discovery pushes back the fossil record of cephalopods by 30 million years, revealing their early evolution.

Area of Science:

  • Paleontology
  • Evolutionary Biology
  • Marine Biology

Background:

  • Burgess Shale deposits preserve early animal body plans.
  • Early cephalopod evolution, particularly nautiloids, is poorly understood.
  • Nectocaris pteryx is a problematic fossil from the Middle Cambrian.

Purpose of the Study:

  • Reinterpret Nectocaris pteryx as a primitive cephalopod.
  • Investigate the early evolution of cephalopods and their body plans.
  • Extend the fossil record of cephalopods into the Cambrian.

Main Methods:

  • Analysis of new fossil material of Nectocaris pteryx from the Burgess Shale.
  • Comparative morphology of Nectocaris and other early Cambrian taxa.
  • Phylogenetic analysis to determine the placement of Nectocarididae within cephalopods.

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Main Results:

  • Nectocaris pteryx is reinterpreted as a stem-group, non-mineralized cephalopod.
  • Nectocaris, Petalilium, and Vetustovermis form the clade Nectocarididae.
  • Primitive cephalopods lacked shells, were hyperbenthic, and possessed a funnel for jet propulsion.

Conclusions:

  • Cephalopod fossil record extended by over 30 million years into the Cambrian.
  • Jet propulsion evolved prior to the acquisition of a mineralized shell in cephalopods.
  • The Cambrian period likely harbored a 'fuse' for the Ordovician diversification of shell-bearing cephalopods.