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Updated: May 27, 2026

3D Culturing of Organoids from the Intestinal Villi Epithelium Undergoing Dedifferentiation
Published on: April 1, 2021
Norifumi Takeda1, Rajan Jain, Matthew R LeBoeuf
1Department of Cell and Developmental Biology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA 19104, USA.
Researchers identified Hopx as a marker for +4 intestinal stem cells. These +4 cells can generate crypt base columnar (CBC) cells, and CBCs can generate +4 cells, revealing a bidirectional relationship between stem cell types.
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