Related Experiment Video
Updated: May 10, 2026

04:28
Eyestalk Ablation to Increase Ovarian Maturation in Mud Crabs
Published on: March 31, 2023
Archaeological shellfish size and later human evolution in Africa.
Richard G Klein1, Teresa E Steele
1Department of Biology, Stanford University, Stanford, CA 94305, USA.
Summary
Early human expansion from Africa (Middle Stone Age) showed no signs of population growth, contrary to previous assumptions linking behavioral changes to increased populations. Archaeological evidence suggests small, stable populations during this period.
Area of Science:
- Archaeology
- Paleoanthropology
- Human Evolution
Background:
- The African Middle Stone Age (MSA) (~200-50 ka) is investigated for the origins of behavioral modernity preceding the global expansion of modern humans.
- Potential early art objects like incised ocher and shell beads from MSA sites suggest cognitive modernity.
Observation:
- Archaeologists often link the emergence of art and behavioral changes to population growth.
- Gastropod shell size in coastal middens can indicate collection intensity, reflecting population size.
Findings:
- Economically important Cape turban shells and limpets from South African MSA layers are significantly larger than those from later Later Stone Age (LSA) layers under similar environmental conditions.
- Larger shell size in MSA sites suggests less intense collection, implying smaller, stable human populations.
Implications:
- Cognitive modernity during the MSA was not driven by population growth.
- Significant human population increase occurred later, near the MSA/LSA transition, coinciding with the Out-of-Africa expansion.
Related Concept Videos
The Fossil Record
The fossil record documents only a small fraction of all organisms that have ever inhabited Earth. Fossilization is a rare process, and most organisms never become fossils. Moreover, the fossil record only exhibits fossils that have been discovered. Nevertheless, sedimentary rock fossils of long-lived, abundant, hard-bodied organisms dominate the fossil record. These fossils offer valuable information, such as an organism's physical form, behavior, and age. Studying the fossil record helps...
What is Evolutionary History?
Scientists record evolutionary history by analyzing fossil, morphological, and genetic data. The fossil record documents the history of life on Earth and provides evidence for evolution. However, both fossil and living organisms offer evidence that outlines Earth’s evolutionary history.
The Evidence for Evolution
Genetic variations accumulating within populations over generations give rise to biological evolution. Evolutionary changes can result in the formation of novel varieties and entire new species. These changes are responsible for the diverse forms of life inhabiting the planet. The evidence for evolution suggests that all living organisms descended from common ancestors.
Speciation Rates
Overview
Optimal Foraging
How animals obtain and eat their food is called foraging behavior. Foraging can include searching for plants and hunting for prey and depends on the species and environment.
Genome Size and the Evolution of New Genes
While every living organism has a genome of some kind (be it RNA, or DNA), there is considerable variation in the sizes of these blueprints. One major factor that impacts genome size is whether the organism is prokaryotic or eukaryotic. In prokaryotes, the genome contains little to no non-coding sequence, such that genes are tightly clustered in groups or operons sequentially along the chromosome. Conversely, the genes in eukaryotes are punctuated by long stretches of non-coding sequence.

