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Updated: May 6, 2026

Optimized Bone Sampling Protocols for the Retrieval of Ancient DNA from Archaeological Remains
Published on: November 30, 2021
Neanderthal axial and appendicular remains from Moula-Guercy, Ardèche, France
Ben Mersey1, Kyle Brudvik, Michael T Black
1Human Evolution Research Center, University of California Berkeley, 3101 Valley Life Sciences Building, Berkeley, CA, 94720-3160; Department of Integrative Biology, University of California Berkeley, 3040 Valley Life Sciences Building, Berkeley, CA, 94720-3140.
Abstract:
Excavations carried out during the 1990s at Moula-Guercy cave Ardèche, France, yielded 108 hominid specimens dating to 100-120 Ka. In this paper, we describe and compare the 39 axial and appendicular specimens not including hand and foot bones. Among these remains are a large adult femur, several clavicles, a likely antimeric pair of radial heads, and a nearly complete superior pubic ramus. Analyses of this material indicate a clear affinity with Neanderthals by the presence of large and robust muscle attachments, thick long bone cortices, a long pubic ramus, and a superoinferiorly flattened clavicle shaft. The recovered remains reveal the presence of a mature male, a smaller mature individual, possibly a reproductive age female, an immature individual of age 10-12, and a second immature individual of age 4. Future analyses on the Moula-Guercy remains will illuminate ties to other known Neanderthal populations and contribute to the ongoing debate over the relative rate of Neanderthal metric growth.
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