Dependence of scroll-wave dynamics on the orientation of a gradient of excitability

The dynamics of scroll waves with a variable orientation to a vertically oriented gradient of excitability is studied by optical tomography in the ferroin-catalyzed Belousov-Zhabotinsky reaction. An almost perpendicular orientation between the scroll wave and gradient induces a pair of twists of opposite handedness on the scroll wave. The position of the nodal plane formed between the twists is governed by the time delay of the twist formation and therefore leads to a symmetric or asymmetric twisted scroll wave. Larger inclinations between scroll wave and gradient cause a drift of the filament along the reactor wall until it reaches the bottom of the reaction container. In this case, the scroll wave does not twist, suggesting that a drift acts as an alternative mechanism of responding to the gradient.