Related Experiment Video
Updated: Mar 31, 2026

Behavioral Assessment of Manual Dexterity in Non-Human Primates
Published on: November 11, 2011
Variability in an early hominin percussive tradition: the Acheulean versus cultural variation in modern chimpanzee
1Archaeology, Classics and Egyptology, School of Histories, Language and Cultures, University of Liverpool, Liverpool L69 3BX, UK gowlett@liverpool.ac.uk.
Abstract:
Percussion makes a vital link between the activities of early human ancestors and other animals in tool-use and tool-making. Far more of the early human actions are preserved as archaeology, since the percussion was largely used for making hard tools of stone, rather than for direct access to food. Both primate tools and early hominin tools, however, offer a means to exploring variability in material culture, a strong focus of interest in recent primate studies. This paper charts such variability in the Acheulean, the longest-lasting tool tradition, extant form about 1.7 to about 0.1 Ma, and well known for its characteristic handaxes. The paper concentrates on the African record, although the Acheulean was also known in Europe and Asia. It uses principal components and discriminant analysis to examine the measurements from 66 assemblages (whole toolkits), and from 18 sets of handaxes. Its review of evidence confirms that there is deep-seated pattern in the variation, with variability within a site complex often matching or exceeding that between sites far distant in space and time. Current techniques of study allow comparisons of handaxes far more easily than for other components, stressing a need to develop common practice in measurement and analysis. The data suggest, however, that a higher proportion of traits recurs widely in Acheulean toolkits than in the chimpanzee record.
Related Concept Videos
Convergent Evolution
The Fossil Record
Evolutionary Psychology
Synteny and Evolution
Around 80 million years ago, the human and mice lineages diverged from the common ancestor. During the course of evolution, the ancestral...
Variation
When independent and dependent variables are plotted on a scatter plot, the slope of a line is a value that describes the rate of change between the two...
Cultural Influences on Personality

