Quantitative measurement of food selection : A modification of the forage ratio and Ivlev's electivity index

The forage ratio and Ivlev's electivity index are common measures to quantify food selection but the values of both indices depend not only on the extent of selection but also on the relative abundances of the food types in the environment. They are therefore useless when food types with different relative abundances are compared, or when the relation between selection and relative abundance is studied. Modified versions of both indices are proposed which are based directly on the rates of decrement (mortality) of the food due to feeding, and are independent of the relative abundance.