Principles of Classical Conditioning
Classical Conditioning
Timing and Consequences on Behavior
Behaviorism
Instinctive Drift
Generalization, Discrimination, and Extinction
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Updated: Feb 16, 2026

Recording Single Neurons' Action Potentials from Freely Moving Pigeons Across Three Stages of Learning
Published on: June 2, 2014
Matthew S Matell1, Rebecca B Della Valle1
1Department of Psychological and Brain Sciences, Villanova University, Villanova, Pennsylvania 19085, USA.
Pavlovian-to-instrumental transfer shows that conditioned stimuli alter operant responding rates. This study reveals that the timing of conditioned stimuli influences response rates, reflecting learned temporal expectancies.
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