The Fossil Record
Synteny and Evolution
What is Evolutionary History?
Lateralization
Convergent Evolution
Bones of the Upper Limb: Humerus
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Updated: Feb 16, 2026

Optimized Bone Sampling Protocols for the Retrieval of Ancient DNA from Archaeological Remains
Published on: November 30, 2021
Marina Lozano1, Almudena Estalrrich2, Luca Bondioli3
1Institut Català de Paleoecologia Humana i Evolució Social (IPHES) and University Rovira i Virgili (URV), Tarragona, Spain.
Fossil hominin teeth reveal a consistent 9:1 right- to left-handedness ratio, indicating modern handedness patterns existed long before Homo sapiens. This ancient trait is evident in early African and European hominin fossils.
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