CRISPR/Cas9 Genome Editing
Mutations
Mutations
Genomic Imprinting and Inheritance
Viral Mutations
Genome Size and the Evolution of New Genes
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Updated: Feb 13, 2026

Using Sniper-Cas9 to Minimize Off-target Effects of CRISPR-Cas9 Without the Loss of On-target Activity Via Directed Evolution
Published on: February 26, 2019
Alejandro Chavez1,2,3, Benjamin W Pruitt2, Marcelle Tuttle2
1Department of Pathology and Cell Biology, Columbia University College of Physicians and Surgeons, New York, NY 10032; ac4304@cumc.columbia.edu gchurch@genetics.med.harvard.edu.
Scientists developed a guide RNA tuning method for Cas9 gene editing to prevent specific mutations. This in vivo system successfully restricted unwanted mutations in engineered organisms, even in the mouse gut.
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