Related Experiment Video
Updated: Jan 4, 2026

Use of Chironomidae Diptera Surface-Floating Pupal Exuviae as a Rapid Bioassessment Protocol for Water Bodies
Published on: July 24, 2015
Phylogenetic Classification of the Atlides Section of the Eumaeini (Lepidoptera, Lycaenidae)
Ananda Regina P Martins1, Marcelo Duarte, Robert K Robbins
1Museu de Zoologia, Universidade de São Paulo, Avenida Nazaré 481, Ipiranga, 04263-000, São Paulo, SP, Brazil. Department of Biology, McGill University, Redpath Museum, 859 Sherbrooke Street West, Montreal, Québec, Zip code: H3A2K6, Canada. ananda.pereiramartins@mail.mcgill.ca.
Abstract:
This paper provides the first phylogenetic classification of the Atlides Section (Lycaenidae: Theclinae, Eumaeini). It is based on a recently published morphological phylogenetic study, in which the Atlides Section proved to be monophyletic in all analyses. In particular, that study identified a hindwing cleft anal lobe in all members of the Atlides Section that is lacking in all other Eumaeini (except for some species of Panthiades Hübner). The included genera are Theritas Hübner; Arcas Swainson; Lucilda d'Abrera Bálint; Pseudolycaena Wallengren; Brangas Hübner; Atlides Hübner; and Denivia K. Johnson. Each is characterized by synapomorphies. Margaritheclus Bálint and Dabreras Bálint are new generic synonyms of Lucilda and Brangas, respectively. We recognize 71 species. Distribution and habitat are provided for each, and nomenclature is updated. Atlides centralis Salazar Henao is a new synonym of Atlides browni Constantino, Salazar K. Johnson; Brangas contrastus Bálint is a new synonym of Brangas felderi (Goodson); Denivia grava Bálint, K. Johnson Kroenlein is a new synonym of Denivia deniva (Hewitson); Denivia ponsanota Bálint, K. Johnson Kroenlein is a new synonym of Atlides zava (Hewitson); Denivia striata Bálint, K. Johnson Kroenlein is a new synonym of Atlides zava (Hewitson); Margaritheclus boliboyeri Bálint Wojtusiak is a new synonym of Margaritheclus boliboyerus Bálint Wojtusiak; and Theritas gozmanyi Bálint Wojtusiak is a new synonym of Theritas paupera (C. Felder R. Felder). New combinations are Lucilda margaritacea (Draudt), Lucilda danaus (C. Felder R. Felder), Lucilda dabrerus (Bálint), Lucilda boliboyerus (Bálint Wojtusiak), Denivia curitabaensis (K. Johnson), Denivia augustinula (Goodson), Denivia theocritus (Fabricius), Denivia augustula (W.F. Kirby), Denivia arene (Goodson), Denivia monica (Hewitson), Denivia hemon (Cramer), Denivia phegeus (Hewitson), Denivia acontius (Goodson), Denivia chaluma (Schaus), Denivia viresco (H.H. Druce), Denivia silma (Martins, Faynel Robbins), and Denivia lisus (Stoll).
More Related Videos
Related Concept Videos
Taxonomy
Hierarchy of Taxonomy
The hierarchy that Carolus Linnaeus first...
Microbial Classification System
Diversity of Protists I
Phylogenetic Trees
Phylogeny
Pollination and Flower Structure

