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Updated: Aug 16, 2025

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Published on: May 7, 2016
Grouping behavior in a Triassic marine apex predator.
Neil P Kelley1, Randall B Irmis2, Paige E dePolo3
1Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, 1001 Madison Drive NW, Washington, DC 20560, USA.
Giant ichthyosaurs, Shonisaurus, exhibit ancient grouping behaviors, with fossil evidence suggesting social or reproductive aggregation in the Late Triassic. This indicates marine tetrapod group behavior dates back at least 230 million years.
Area of Science:
- Paleontology
- Marine Biology
- Evolutionary Biology
Background:
- Marine tetrapods play key ecological roles and often form aggregations.
- Grouping behaviors in marine tetrapods are driven by prey, social factors, or oceanography.
Purpose of the Study:
- To investigate early marine tetrapod grouping behaviors.
- To document evidence of aggregation in the giant ichthyosaur Shonisaurus.
Main Methods:
- Analysis of numerous Shonisaurus specimens from the Luning Formation, Nevada.
- Examination of fossil distribution, abundance, and demographic composition.
- Integration of geological evidence from the Late Triassic marine environment.
Main Results:
- A large, monotaxic assemblage of Shonisaurus fossils spanning extensive area and time.
- Disproportionate abundance of large individuals and neonatal/embryonic specimens, lacking intermediate juveniles.
- Evidence suggests Shonisaurus formed dense aggregations in a tropical marine environment for millennia.
Conclusions:
- Philopatric grouping behavior in marine tetrapods is at least 230 million years old.
- Shonisaurus aggregations were potentially linked to reproductive activities.
- This study reveals ancient social behaviors in early marine vertebrates.
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