Related Experiment Video
Updated: May 28, 2025

Daily Transfers, Archiving Populations, and Measuring Fitness in the Long-Term Evolution Experiment with Escherichia coli
Published on: August 18, 2023
A reassessment of the "hard-steps" model for the evolution of intelligent life
Daniel B Mills1,2,3, Jennifer L Macalady2,3,4,5, Adam Frank6
1Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universität München, 80333 Munich, Germany.
Abstract:
According to the "hard-steps" model, the origin of humanity required "successful passage through a number of intermediate steps" (so-called "hard steps") that were intrinsically improbable in the time available for biological evolution on Earth. This model similarly predicts that technological life analogous to human life on Earth is "exceedingly rare" in the Universe. Here, we critically reevaluate core assumptions of the hard-steps model through the lens of historical geobiology. Specifically, we propose an alternative model where there are no hard steps, and evolutionary singularities required for human origins can be explained via mechanisms outside of intrinsic improbability. Furthermore, if Earth's surface environment was initially inhospitable not only to human life, but also to certain key intermediate steps required for human existence, then the timing of human origins was controlled by the sequential opening of new global environmental windows of habitability over Earth history.
Related Concept Videos
Evolutionary Psychology
What is Evolutionary History?
The Evidence for Evolution
Biological Influences on Intelligence
Environmental Influences on Intelligence
Limits to Natural Selection

