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Crystal structure of the human class II MHC protein HLA-DR1 complexed with an influenza virus peptide
L J Stern1, J H Brown, T S Jardetzky
1Department of Biochemistry and Molecular Biology, Harvard University, Cambridge Massachusetts 62138.
Abstract:
An influenza virus peptide binds to HLA-DR1 in an extended conformation with a pronounced twist. Thirty-five per cent of the peptide surface is accessible to solvent and potentially available for interaction with the antigen receptor on T cells. Pockets in the peptide-binding site accommodate five of the thirteen side chains of the bound peptide, and explain the peptide specificity of HLA-DR1. Twelve hydrogen bonds between conserved HLA-DR1 residues and the main chain of the peptide provide a universal mode of peptide binding, distinct from the strategy used by class I histocompatibility proteins.
Insights
This study reveals how an influenza peptide binds to HLA-DR1, showing a unique twist and specific pocket interactions. This binding mode, with key hydrogen bonds, differs from class I histocompatibility proteins.
Area of Science:
- Immunology
- Structural Biology
- Virology
Background:
- Human Leukocyte Antigen (HLA) molecules present peptides to T cells, crucial for immune response.
- Understanding peptide-HLA interactions is key to vaccine development and autoimmune disease research.
Purpose of the Study:
- To elucidate the structural basis of peptide binding to HLA-DR1.
- To characterize the conformation and interaction details of an influenza virus peptide within the HLA-DR1 binding site.
Main Methods:
- X-ray crystallography or cryo-EM to determine the structure of the peptide-HLA-DR1 complex.
- Computational analysis to assess peptide surface accessibility and binding pocket interactions.
Main Results:
- The influenza peptide adopts an extended conformation with a significant twist when bound to HLA-DR1.
- 35% of the peptide surface is solvent-accessible, available for T cell receptor interaction.
- Specific pockets within HLA-DR1 accommodate five peptide side chains, dictating specificity.
- Twelve hydrogen bonds between conserved HLA-DR1 residues and the peptide backbone establish a conserved binding mode.
Conclusions:
- The binding of influenza peptides to HLA-DR1 involves a unique, twisted conformation and specific side-chain interactions.
- This binding mechanism is distinct from that observed in class I histocompatibility proteins, highlighting diverse peptide presentation strategies.