1Department of Biochemistry and Biophysics, University of Pennsylvania, Philadelphia 19104-6059, USA. sharp@crystal.med.upenn.edu
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Mutating hirudin's acidic residues (aspartate and glutamate) to asparagine or glutamine reduced anticoagulant activity by altering electrostatic interactions with thrombin. Structural changes can compensate for unfavorable interactions in multiple mutants.
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