1Nebraska Behavioural Biology Group, School of Biological Sciences, University of Nebraska, Lincoln, NE 68588, USA. strixkn@fish.fish.kokudai.ac.jp
Finite population size and payoff variance in the iterated Prisoner's Dilemma (IPD) challenge standard ESS analysis. Tit-for-tat (TFT) strategies face invasion risks from all-defect strategies (ALLD) due to payoff variance, not just mean payoffs.
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