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IGF-I induces collagen and IGFBP-5 mRNA in rat intestinal smooth muscle

E M Zimmermann1, L Li, Y T Hou

  • 1Department of Internal Medicine, University of Michigan Medical School, Ann Arbor 48109-0586, USA.

Insights

Insulin-like growth factor binding protein 5 (IGFBP-5) mRNA increases in inflamed rat intestines, particularly in smooth muscle. Insulin-like growth factor-I (IGF-I) boosts IGFBP-5 and collagen mRNA in these cells.

Area of Science:

  • Gastroenterology
  • Molecular Biology
  • Cell Biology

Background:

  • Intestinal inflammation, such as peptidoglycan-polysaccharide enterocolitis, can alter gene expression in the gut.
  • Insulin-like growth factor (IGF) binding proteins play crucial roles in regulating IGF bioavailability and function.
  • IGF binding protein 5 (IGFBP-5) is implicated in various cellular processes, including cell growth and extracellular matrix production.

Purpose of the Study:

  • To investigate the expression and regulation of IGFBP-5 mRNA in the rat intestine during enterocolitis.
  • To determine the effect of IGF-I on IGFBP-5 and collagen type I mRNA synthesis in cultured intestinal smooth muscle cells.

Main Methods:

  • Northern analysis and in situ hybridization were used to quantify IGFBP-5 mRNA levels in rat colon tissue.
  • Primary rat intestinal smooth muscle cells were cultured to assess the impact of IGF-I on gene expression.
  • Western ligand blot and immunoblotting were employed to measure IGFBP-5 protein levels.
  • Transcriptional blockade and cycloheximide treatment were used to investigate mRNA stability and protein synthesis dependence.

Main Results:

  • IGFBP-5 mRNA levels were significantly elevated (2.4-fold) in the inflamed colon of rats compared to controls.
  • IGFBP-5 mRNA showed high expression in the smooth muscle layer of the rat intestine.
  • In cultured smooth muscle cells, IGF-I treatment increased IGFBP-5 mRNA (1.9-fold) and collagen type alpha1(I) mRNA (1.6-fold).
  • IGF-I also dose- and time-dependently increased secreted IGFBP-5 protein levels, without affecting mRNA decay rate.

Conclusions:

  • IGFBP-5 mRNA is expressed in intestinal smooth muscle cells and its expression is upregulated during chronic intestinal inflammation.
  • IGF-I stimulates the synthesis of IGFBP-5 and collagen type I in intestinal smooth muscle cells, suggesting a role in inflammation-associated tissue remodeling.

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