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Molecular Biology of the Cell|June 11, 2026
The ESCRT-0 protein HRS regulates hepatocellular lipid droplet catabolismMathilda M Willoughby, Ankit Shroff, Bridget E Crossman, et al.Biochimica Et Biophysica Acta. Molecular Cell Research|October 6, 2020
One ring to bring them all and in the darkness bind them: The trafficking of heme without deliverersIan G Chambers, Mathilda M Willoughby, Iqbal Hamza, et al.Biorxiv : the Preprint Server for Biology|November 26, 2025
The ESCRT-0 protein HRS regulates hepatocellular lipid droplet catabolismMathilda M Willoughby, Ankit Shroff, Bridget E Crossman, et al.The American Journal of Pathology|August 28, 2025
Lipid Droplet Dynamics in Alcoholic SteatohepatitisDelia Y Omar, Mathilda M Willoughby, Nourhan Mostafa, et al.Journal of Cell Science|April 9, 2020
Mitochondrial-nuclear heme trafficking in budding yeast is regulated by GTPases that control mitochondrial dynamics and ER contact sitesOsiris Martinez-Guzman, Mathilda M Willoughby, Arushi Saini, et al.Methods in Molecular Biology (Clifton, N.J.)|July 15, 2024
Fluorometric Methods to Measure Bioavailable and Total HemeIramofu M Dominic, Mathilda M Willoughby, Abigail K Freer, et al.Nature|October 19, 2022
HRG-9 homologues regulate haem trafficking from haem-enriched compartmentsFengxiu Sun, Zhenzhen Zhao, Mathilda M Willoughby, et al.Redox Biology|September 13, 2021
Mitochondrial contact site and cristae organizing system (MICOS) machinery supports heme biosynthesis by enabling optimal performance of ferrochelataseJonathan V Dietz, Mathilda M Willoughby, Robert B Piel, et al.Journal of Inorganic Biochemistry|October 23, 2025
Progesterone receptor membrane component 1 (PGRMC1) regulates Heme trafficking through mitochondria-ER junctionsRobert B Piel Iii, Chibuike D Obi, Martonio Ponte Viana, et al.Biorxiv : the Preprint Server for Biology|May 20, 2024
The molecular mechanism of on-demand sterol biosynthesis at organelle contact sitesNaama Zung, Nitya Aravindan, Angela Boshnakovska, et al.Pageof 1