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Updated: Aug 9, 2026

Behavioral Assessment of Manual Dexterity in Non-Human Primates
Published on: November 11, 2011
DEVELOPMENT OF A COMPLEX MULTIPLE SCHEDULE IN THE CHIMPANZEE
Chimpanzee behavior was studied using a complex multiple schedule involving avoidance and reinforcement tasks. This research informed the space flights of chimpanzees Ham and Enos.
Area of Science:
- Behavioral science
- Primatology
- Space biology
Background:
- The study details the development of chimpanzee behavior under a complex four-component, three-lever multiple schedule.
- This schedule incorporated various operant conditioning procedures, including avoidance and reinforcement tasks.
Purpose of the Study:
- To describe and analyze chimpanzee behavior under a sophisticated multiple schedule.
- To assess the applicability of these behavioral paradigms in the context of space exploration.
Main Methods:
- A four-component, three-lever multiple schedule was designed and implemented.
- Component schedules included Sidman avoidance, discriminated avoidance, fixed ratio, differential reinforcement of low rate, and symbol discrimination tasks.
Main Results:
- The avoidance component was utilized in the suborbital space flight of chimpanzee Ham on January 31, 1961.
- Chimpanzee Enos performed on the multiple schedule during a three-orbit space flight on November 29, 1961.
Conclusions:
- The study demonstrated the feasibility of using complex behavioral schedules with chimpanzees in spaceflight.
- This research provided valuable data on primate behavior under extreme environmental conditions.
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