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Updated: May 19, 2026

Behavioral Assessment of Manual Dexterity in Non-Human Primates
Published on: November 11, 2011
Stone tool production and utilization by bonobo-chimpanzees (Pan paniscus)
Itai Roffman1, Sue Savage-Rumbaugh, Elizabeth Rubert-Pugh
1International Graduate Center of Evolution, Institute of Evolution, University of Haifa, Mount Carmel, Haifa 31905, Israel.
Abstract:
Using direct percussion, language-competent bonobo-chimpanzees Kanzi and Pan-Banisha produced a significantly wider variety of flint tool types than hitherto reported, and used them task-specifically to break wooden logs or to dig underground for food retrieval. For log breaking, small flakes were rotated drill-like or used as scrapers, whereas thick cortical flakes were used as axes or wedges, leaving consistent wear patterns along the glued slits, the weakest areas of the log. For digging underground, a variety of modified stone tools, as well as unmodified flint nodules, were used as shovels. Such tool production and utilization competencies reported here in Pan indicate that present-day Pan exhibits Homo-like technological competencies.
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Quarrying of Stone
One common method involves using a diamond belt saw to cut large blocks from the quarry face. These blocks can be about 50 feet long and 12 feet high. After the initial vertical cut, drilling is performed at the base of the block.
The Fossil Record

