The Fossil Record
What is Evolutionary History?
The Colonization of Land
Convergent Evolution
Bone Formation by Intramembranous Ossification
Phylogeny
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Updated: Mar 17, 2026

Processing Embryo, Eggshell, and Fungal Culture for Scanning Electron Microscopy
Published on: August 16, 2019
Tyler R Lyson1, Bruce S Rubidge2, Torsten M Scheyer3
1Department of Earth Sciences, Denver Museum of Nature and Science, Denver, CO 80205, USA; Evolutionary Studies Institute, University of the Witwatersrand, PO Box Wits, 2050 Johannesburg, South Africa.
The evolution of the turtle shell began with broadened ribs, initially for digging (fossoriality), not protection. This adaptation in early turtles like Eunotosaurus influenced their body plan and survival.
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