The Spindle Assembly Checkpoint
Meiosis II
Spindle Assembly
Separation of Sister Chromatids
Anaphase A and B
The Mitotic Spindle
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Updated: Mar 1, 2026

Evaluation of the Spindle Assembly Checkpoint Integrity in Mouse Oocytes
Published on: September 13, 2022
Prajakta Varadkar1, Fatima Abbasi1, Kazuyo Takeda1
1a Division of Cellular and Gene Therapies, Center for Biologics Evaluation and Research , US Food and Drug Administration , Silver Spring , MD.
The Spindle Assembly Checkpoint (SAC) requires PP2A-B56γ to stabilize BubR1, preventing premature cell division. Loss of PP2A-B56γ leads to chromosomal instability, a cancer hallmark.
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