[Shade tolerance and the adaptability of forest plants in morphology and physiology: A review.]

Shade tolerance, defined as the ability to survive and grow under low light, plays an important role in the succession of forest plant communities. The mechanism of adaptation of plants to shading has become an important topic in ecology research. In this paper, we reviewed shade tole-rance of various forest plants and examined their adaptability in terms of morphology and physiology, and discussed the plasticity on growth traits, biomass distribution, crown structure, leaf morphology and physiology, leaf anatomical structure, photosynthetic parameters, carbohydrate allocation, water and nutrient utilization related to shade tolerance of forest plants. Finally, we discussed some problems with previous studies, and proposed possible future research issues.