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Related Concept Videos

Anatomy of the Ear01:16

Anatomy of the Ear

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Auditory sensation, commonly called hearing, involves the transformation of sonic waves into neural impulses facilitated by the structures of the auditory organ. The prominent, flesh-like structure on the side of the head, called the auricle, directs sound waves towards the auditory canal. The auricle is often mislabeled as the pinna, a term more aligned with mobile structures like a feline's external ear. The auditory canal penetrates the cranium via the external auditory meatus of the...
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Hearing01:31

Hearing

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When we hear a sound, our nervous system is detecting sound waves—pressure waves of mechanical energy traveling through a medium. The frequency of the wave is perceived as pitch, while the amplitude is perceived as loudness.
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Auditory Pathway01:15

Auditory Pathway

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Auditory pathways constitute the complex neural circuits responsible for transmitting and interpreting auditory information from the peripheral auditory system to the brain. Sound waves are initially captured by the outer ear, funneled through the ear canal, and reach the tympanic membrane (eardrum). These vibrations are transmitted via the middle ear's ossicles to the inner ear's cochlea.
When viewed cross-sectionally, the cochlea reveals the scala vestibuli and scala tympani flanking...
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Assessing Body Temperature - Tympanic membrane01:14

Assessing Body Temperature - Tympanic membrane

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Assessing tympanic membrane temperature involves using a tympanic membrane thermometer (TMT). Here is a step-by-step guide:
Step 1: Begin by practicing good hand hygiene to prevent the transmission of microorganisms.
Step 2: Turn on the thermometer and wait until the ready sign appears on the screen to ensure accurate measurement.
Step 3: Slide the probe cover in place to prevent cross-contamination.
Step 4: Instruct the patient to tilt their head to the side for comfort and check for cerumen...
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Auditory Perception01:17

Auditory Perception

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The auditory system is essential for sound perception, utilizing various critical structures. When sound waves enter the outer ear, they travel through the ear canal and cause the eardrum to vibrate. These vibrations are then transmitted to the middle ear, where three tiny bones – the malleus, incus, and stapes – amplify the sound. This amplification is crucial, as it ensures that the sound vibrations are strong enough to be conveyed to the inner ear. These vibrations then reach the...
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The Auditory Ossicles01:11

The Auditory Ossicles

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The auditory ossicles of the middle ear transmit sounds from the air as vibrations to the fluid-filled cochlea. The auditory ossicles consist of two malleus (hammer) bones, two incus (anvil) bones, and two stapes (stirrups), one on each side. These bones develop during the fetal stage and are the ones to ossify first. They are fully mature at birth and do not grow afterward.
The aptly named stapes look very much like a stirrup. The three ossicles are unique to mammals, and each plays a role in...
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Inhibitory inputs to avian ITD circuits.

Trends in hearing·2026
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A general mechanism of airborne hearing in recent and early non-tympanate tetrapods.

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Vomeronasal organ of the North American river otter (Lontra canadensis): Morphological and evolutionary insights based on iodine-enhanced computed tomography.

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Related Experiment Video

Updated: Sep 7, 2025

Discovering Middle Ear Anatomy by Transcanal Endoscopic Ear Surgery: A Dissection Manual
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Discovering Middle Ear Anatomy by Transcanal Endoscopic Ear Surgery: A Dissection Manual

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Hearing without a tympanic ear.

Grace Capshaw1, Jakob Christensen-Dalsgaard2, Catherine E Carr1

  • 1Department of Biology, University of Maryland, College Park, MD 20742, USA.

The Journal of Experimental Biology
|June 20, 2022
PubMed
Summary
This summary is machine-generated.

Hearing evolved in land vertebrates before the development of the tympanic middle ear. Recent studies reveal sound detection and directional hearing mechanisms in

Keywords:
AuditoryBone conductionExtratympanicSoundVibration

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Area of Science:

  • Evolutionary biology
  • Bioacoustics
  • Vertebrate audiology

Background:

  • The evolution of sensitive hearing in terrestrial vertebrates is marked by a significant gap between early tetrapods and the emergence of the tympanic middle ear.
  • Understanding the auditory system's ancestral state is crucial for deciphering the evolution of hearing on land.

Purpose of the Study:

  • To investigate the mechanisms of hearing in extant atympanate (lacking a tympanic membrane) vertebrates.
  • To propose a generalized model for sound detection and directional hearing in early tetrapods and other 'earless' vertebrates.
  • To review the evolutionary history of the tympanic middle ear, including its multiple origins and losses.

Main Methods:

  • Review of extant atympanate vertebrate hearing mechanisms.
  • Analysis of extratympanic sound transmission pathways.
  • Synthesis of studies on tympanate and atympanate hearing, including infrasound reception and human bone conduction.

Main Results:

  • Atympanate vertebrates possess functional mechanisms for sound pressure detection and directional hearing.
  • Extratympanic pathways play a significant role in transmitting sound to the inner ear.
  • The tympanic middle ear has a complex evolutionary history with multiple independent acquisitions and losses.

Conclusions:

  • Hearing in early tetrapods likely involved extratympanic mechanisms, predating the evolution of the tympanic middle ear.
  • These findings provide insights into the ancestral auditory capabilities of terrestrial vertebrates.
  • Understanding diverse auditory mechanisms, including extratympanic pathways, is essential for a comprehensive view of hearing evolution.