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    Ecosystems connected by resource flows form networks. Network structure, not just flow rates, significantly impacts landscape-scale ecosystem function, influenced by organism traits and resource availability.

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    Area of Science:

    • Ecology
    • Environmental Science
    • Network Science

    Background:

    • Ecosystems are interconnected through the flow of nonliving resources.
    • Previous studies focused on few coupled ecosystems, neglecting broader network structures.
    • Understanding resource flow networks is crucial for metaecosystem function.

    Purpose of the Study:

    • Investigate how the spatial structure of resource flow networks influences metaecosystem function.
    • Analyze the impact of homogeneous versus heterogeneous network distributions.
    • Determine the interplay between network structure, biotic, and abiotic factors.

    Main Methods:

    • Utilized metaecosystem models with varying resource flow network structures.
    • Compared outcomes between homogeneously and heterogeneously distributed networks.
    • Analyzed how local nonlinear processes scale up within the network.

    Main Results:

    • Metaecosystem function differs significantly based on resource flow network structure.
    • Network structure influences spatial variation in local dynamics, affecting overall function.
    • Optimal function depends on the interaction between network structure, organism traits, and resource flow rates.

    Conclusions:

    • Spatial structure of resource flow networks is a key driver of landscape-scale ecosystem function.
    • Modifications to network structure, biotic, or abiotic properties have substantial effects on metaecosystem function.
    • Future research should consider network topology in ecosystem management and conservation.