Related Experiment Video
Updated: Jul 12, 2026

05:30
Technique for Studying Arthropod and Microbial Communities within Tree Tissues
Published on: November 16, 2014
Summary
Fossil fern phloem from the Pennsylvanian period reveals sieve elements with pores on radial walls. This discovery aids in comparing ancient fern vascular tissue with that of modern species.
Area of Science:
- Paleobotany
- Plant anatomy
- Evolutionary biology
Background:
- The Pennsylvanian period (approx. 323-299 million years ago) was a critical time for early vascular plant evolution.
- Understanding ancient plant vascular systems, particularly phloem, is key to tracing plant evolution.
- The fern genus Etapteris is known from fossil records, but its phloem structure requires detailed investigation.
Purpose of the Study:
- To describe the detailed structure of phloem tissue in the Pennsylvanian fern Etapteris.
- To identify and characterize sieve elements and associated structures within Etapteris phloem.
- To establish comparative anatomical links between fossil and extant fern phloem.
Main Methods:
- Analysis of permineralized specimens of the fern Etapteris.
- Microscopic examination of phloem tissue, focusing on sieve elements.
- Detailed morphological description of sieve areas and pore distribution.
Main Results:
- Permineralized specimens of Etapteris clearly exhibit phloem tissue.
- Sieve elements possess distinct sieve areas located on their radial walls.
- These sieve areas contain regularly aligned pores, indicating functional vascular connections.
Conclusions:
- The phloem structure of Etapteris provides significant anatomical data for understanding early fern vascular systems.
- The presence of specific sieve element features allows for direct comparison with the phloem of extant ferns.
- This study enhances our knowledge of plant evolution during the Pennsylvanian period through detailed fossil analysis.
Related Concept Videos
Phloem and Sugar Transport
Like many living organisms, plants have tissues that specialize in specific plant functions. For example, shoots are well adapted to rapid growth, while roots are structured to acquire resources efficiently. However, sugar production is primarily restricted to the photosynthetic cells that reside in the leaves of angiosperm plants. Sugar and other resources are transported from photosynthetic tissues to other specialized tissues by a process called translocation.
Plant Tissues
Plants are multicellular eukaryotes with tissue systems made of various cell types that carry out specific functions. Different tissues work together to perform a unique function and form an organ. Organs working together form organ systems. Vascular plants have two distinct organ systems: a shoot system and a root system. The shoot system consists of two portions: the vegetative (non-reproductive) parts of the plant, such as the leaves and the stems, and the reproductive parts of the plant,...
Seedless Vascular Plants
Seedless Vascular Plants Were the First Tall Plants on Earth
The Apoplast and Symplast
Plant growth depends on its ability to take up water and dissolved minerals from the soil. The root system of every plant is equipped with the necessary tissues to facilitate the entry of water and solutes. The plant tissues involved in the transport of water and minerals have two major compartments - the apoplast and the symplast. The apoplast includes everything outside the plasma membrane of living cells and consists of cell walls, extracellular spaces, xylem, phloem, and tracheids. The...
Xylem and Transpiration-driven Transport of Resources
The xylem of vascular plants distributes water and dissolved minerals that are taken up by the roots to the rest of the plant. The cells that transport xylem sap are dead upon maturity, and the movement of xylem sap is a passive process.
Water and Mineral Acquisition
Specialized tissues in plant roots have evolved to capture water, minerals, and some ions from the soil. Roots exhibit a variety of branching patterns that facilitate this process. The outermost root cells have specialized structures called root hairs that increase the root surface, thus increasing soil contact. Water can passively cross into roots, as the concentration of water in the soil is higher than that of the root tissue. Minerals, in contrast, are actively transported into root cells.

