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Updated: Jul 5, 2026

Determining Basal Energy Expenditure and the Capacity of Thermogenic Adipocytes to Expend Energy in Obese Mice
Published on: November 11, 2021
Energy balance during backpacking
L C Hill1, D P Swain, E L Hill
1Health, Physical Education and Exercise Science, Norfolk State University, Norfolk, Virginia 23504, United States. lchill@nsu.edu
Abstract:
Energy expenditure and energy intake were determined in three subjects during a 160-kilometer backpacking trip. Prior to the trip, heart rate and oxygen consumption during treadmill walking while carrying a backpack were measured. Subjects recorded heart rate during hiking. Heart rate was used to estimate energy expenditure based on the heart rate : oxygen consumption relationship (Heart Rate Method). Expenditure was also estimated from distance walked and elevation gained (Terrain Method). Subjects recorded food consumption, and were weighed upon finishing the hike. Mean heart rate during hiking was 105 +/- 12 beats per minute. According to the Heart Rate Method, net expenditure averaged 3410 +/- 955 kilocalories on days 1 - 4, and 2586 +/- 974 kilocalories on day 5. Net expenditure from the Terrain Method was approximately 28 % lower. Gross expenditure, using the Heart Rate Method, averaged 4928 +/- 1050 kilocalories on days 1 - 4, and 3550 +/- 1052 kilocalories on day 5. Energy intake averaged 2134 +/- 625 kilocalories on days 1 - 4, and 1117 +/- 320 kilocalories on day 5. Expected weight loss due to the caloric deficit was 1.8 +/- 0.4 kilograms, and actual weight loss was 1.7 +/- 0.6 kilograms. Subjects expended nearly 5000 kilocalories per day and did not consume sufficient food to maintain body mass.
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